Homeostasis is a term which essentially means "balance" or "equilibrium". Homeostasis is the property of an open system to regulate its internal environment so as to maintain a stable condition, by means of multiple dynamic equilibrium adjustments controlled by interrelated regulation mechanisms. The term was coined in 1932 by Walter Cannon from two Greek words (to remain the same).

Table of contents
1 Biology
2 Feedback
3 Ecological homeostasis
4 Biological homeostasis


The term usually is used in the sense of biological homeostasis. Multicellular organisms require a homeostatic internalenvironment, in order to live; many environmentalists believe this principal also applies to the external environment. Ecological, biological, and social systems are homeostatic. They oppose change to maintain equilibrium. If the system does not succeed in reestablishing its balance, it will be in a state where constraints are more severe than before. This can ultimately lead to the destruction of the system.

Complex systems, such as a human body, must have homeostasis to maintain stability and to survive. These systems do not only have to endure to survive; they must adapt themselves and evolve to modifications of the environment and it must evolve.

Homeostatic systems show several properties

  • they are ultrastable;
  • their whole organisation, internal, structural, and functional, contributes to the maintenance of equilibrium
  • they are unpredictable (the resulting effect of a precise action often has the oppposite effect to what was expected)

A frequent paradox for those responsible for the maintenance and evolution of a complex system (be it a body, an ecosystem, a state, or a planet) is how can a stable organization whose goal is to maintain itself and endure, be able to change and evolve?


When a change of variable is occurring, there are many types of
feedback with which the system to react.

Negative feedback is a reaction in which the system responds in such a way as to reverse the direction of change. Since this tends to keep things constant, it allows the maintenance of homeostasis. For instance, when the concentration of carbon dioxide in the human body increases, the lungs are signalled to increase their activity and expel more carbon dioxide.

In positive feedback, the response is to amplify the change in the variable. This has a de-stabilizing effect, so does not result in homeostasis. Positive feedback is less common in naturally occurring systems than negative feedback, but it has its applications. For example, in nerves, a threshold electric potential triggers the generation of a much larger action potential. (See also leverage points.)

Ecological homeostasis

In the Gaia hypothesis, James Lovelock stated that the entire mass of living matter on Earth (or any planet with life) functions as a vast organism that actively modifies its planet to produce the environment that suits its needs. In this view, the entire planet maintains homeostasis. Whether this sort of system is present on Earth is still open to debate. However, some relatively simple homeostatic mechanisms are generally accepted. For example, when atmospheric carbon dioxide levels rise, plants are able to grow better and thus remove more carbon dioxide from the atmosphere.

Biological homeostasis

Homeostasis is one of the fundamental characteristics of living things. It is the maintenance of the internal environment within tolerable limits.

With regard to any parameter, an organism may be a conformer or a regulator. Regulators try to maintain the parameter at a constant level, regardless of what is happening in its environment. Conformers allow the environment to determine the parameter. For instance, endothermic animals maintain a constant body temperature, while ectothermic animals exhibit wide variation in body temperature.

This is not to say that conformers may not have behavioral adaptations that allow them to exert some control over the parameter in question. For instance, reptiles often sit on sun-heated rockss in the morning to raise their body temperatures.

An advantage of homeostatic regulation is that it allows the organism to function more effectively. For instance, ectotherms tend to become sluggish at low temperatures, whereas endotherms are as active as always. On the other hand, regulation reqires energy. One reason why snakes can eat only once a week is that they use much less energy for maintaining homeostasis.

Homeostasis in the human body

All sorts of factors affect the suitability of our body fluids to sustain life; these include properties like temperature, salinity, acidity (carbon dioxide), and the concentrations of nutrients and wastes (urea, glucose, various ion, oxygen). Since these properties affect the chemical reactions that keep bodies alive, there are built-in physiological mechanisms to maintain them at desirable levels.

This control is achieved with various organs in the body. For example:

  • Thermal regulation
    • The skeletal muscles can shiver to produce heat if the body temperature is too low.
    • Non-shivering thermogenesis involves the decomposition of fat to produce heat.
    • Sweating cools the body with the use of evaporation.
  • Chemical regulation
Most of these organs are controlled by hormones secreted from the pituitary gland, which in turn is directed by the hypothalamus.

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